Red-cockaded Woodpecker
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The red-cockaded woodpecker (''Leuconotopicus borealis'') is a
woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions. ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the southeastern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
.


Description

The red-cockaded woodpecker is small to mid-sized species, being intermediate in size between North America's two most widespread woodpeckers (the
downy Downy, also known as Lenor in Europe, Russia and Japan, is a brand of fabric softener produced by Procter & Gamble that was introduced in 1960. Lenor is a brand name of fabric softener and dryer sheets, also produced by P&G, sold in Europe, Rus ...
and
hairy woodpecker The hairy woodpecker (''Leuconotopicus villosus'') is a medium-sized woodpecker that is found over a large area of North America. It is approximately in length with a wingspan. With an estimated population in 2020 of almost nine million individ ...
s). This species measures in length, spans across the wings and weighs . Among the standard measurements, the wing chord is , the tail is , the bill is and the tarsus is . Its back is barred with black and white horizontal stripes. The red-cockaded woodpecker's most distinguishing feature is a black cap and nape that encircle large white cheek patches. Rarely visible, except perhaps during the breeding season and periods of
territorial A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
defense, the male has a small red streak on each side of its black cap called a ''cockade'', hence its name. The species is listed as Near Threatened by the IUCN and as Endangered by the United States Fish and Wildlife Service.


Behavior

The red-cockaded woodpecker feeds primarily on ants,
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s,
cockroach Cockroaches (or roaches) are a paraphyletic group of insects belonging to Blattodea, containing all members of the group except termites. About 30 cockroach species out of 4,600 are associated with human habitats. Some species are well-known as ...
es,
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s, wood-boring insects, and
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s, and occasionally
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
and
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, rasp ...
. The vast majority of foraging is on pines, with a strong preference for large trees, though they will occasionally forage on
hardwoods Hardwood is wood from dicot trees. These are usually found in broad-leaved temperate and tropical forests. In temperate and boreal latitudes they are mostly deciduous, but in tropics and subtropics mostly evergreen. Hardwood (which comes from ...
and even on corn earworms in cornfields.Jackson, J. A. (1994). Red-cockaded Woodpecker (Dryobates borealis), version 2.0. In The Birds of North America (A. F. Poole and F. B. Gill, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bna.85 Red-cockaded woodpeckers are a territorial, nonmigratory,
cooperative breeding Cooperative breeding is a social system characterized by alloparental care: offspring receive care not only from their parents, but also from additional group members, often called helpers. Cooperative breeding encompasses a wide variety of group s ...
species, frequently having the same mate for several years. The
nesting season The nesting season is the time of year during which birds and some other animals, particularly some reptiles, build nests, lay eggs in them, and in most cases bring up their young. It is usually in the spring. Bird conservation Bird conservat ...
runs from April to June. The breeding female lays three to four
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
s in the breeding male's roost cavity. Group members incubate the small white eggs for 10–13 days. Once hatched, the nestlings remain in the nest cavity for about 26–29 days. Upon fledging, the young often remain with the parents, forming groups of up to nine or more members, but more typically three to four members. There is only one pair of breeding birds within each group, and they normally only raise a single brood each year. The other group members, called helpers, usually males from the previous breeding season, help incubate the eggs and raise the young. Juvenile females generally leave the group before the next breeding season, in search of solitary male groups. The main predators of red-cockaded nests are
rat snake Rat snakes are members – along with kingsnakes, milk snakes, vine snakes and indigo snakes – of the subfamily Colubrinae of the family Colubridae. They are medium to large constrictors and are found throughout much of the Northern Hemi ...
s, although corn snakes also represent a threat.Longleaf Alliance
Studies have also explored the possibility that southern flying squirrels might have a negative impact on red-cockaded woodpecker populations due competition over cavities and predation on eggs and nestlings.


Distribution and habitat

Historically, this woodpecker's range extended in the
southeastern United States The Southeastern United States, also referred to as the American Southeast or simply the Southeast, is a geographical region of the United States. It is located broadly on the eastern portion of the southern United States and the southern por ...
from
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
to
New Jersey New Jersey is a state in the Mid-Atlantic and Northeastern regions of the United States. It is bordered on the north and east by the state of New York; on the east, southeast, and south by the Atlantic Ocean; on the west by the Delaware ...
and
Maryland Maryland ( ) is a state in the Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware and the Atlantic Ocean to ...
, as far west as eastern
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
and
Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...
, and inland to
Missouri Missouri is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it is bordered by eight states (tied for the most with Tennessee ...
,
Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virginia to ...
, and
Tennessee Tennessee ( , ), officially the State of Tennessee, is a landlocked state in the Southeastern region of the United States. Tennessee is the 36th-largest by area and the 15th-most populous of the 50 states. It is bordered by Kentucky to th ...
. Today it is estimated that there are about 5,000 groups of red-cockaded woodpeckers, or 12,500 birds, from Florida to
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...
and west to southeast Oklahoma and eastern Texas, representing about 1% of the woodpecker's original population (over 1 million individuals at one point). They have become locally extinct (
extirpated Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
) in Kentucky, Maryland, Missouri, New Jersey, and Tennessee. The red-cockaded woodpecker makes its home in fire-dependent
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
s.
Longleaf pine The longleaf pine (''Pinus palustris'') is a pine species native to the Southeastern United States, found along the coastal plain from East Texas to southern Virginia, extending into northern and central Florida. In this area it is also known as ...
s (''Pinus palustris'') are most commonly preferred, but other species of southern pine are also acceptable. While other woodpeckers bore out cavities in dead trees where the wood is rotten and soft, the red-cockaded woodpecker is the only one that excavates cavities exclusively in living pine trees. The older pines favored by the red-cockaded woodpecker often suffer from a fungal infection called red heart rot which attacks the center of the trunk, causing the inner wood, the heartwood, to become soft. Cavities are generally excavated over 1 to 3 years. The aggregate of cavity trees is called a cluster and may include 1 to 20 or more cavity trees on 3 to 60
acre The acre is a unit of land area used in the imperial Imperial is that which relates to an empire, emperor, or imperialism. Imperial or The Imperial may also refer to: Places United States * Imperial, California * Imperial, Missouri * Imp ...
s (12,000 to 240,000 m²). The average cluster is about 10 acres (40,000 m²). Cavity trees that are being actively used have numerous, small
resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on natu ...
wells which exude
sap Sap is a fluid transported in xylem cells (vessel elements or tracheids) or phloem sieve tube elements of a plant. These cells transport water and nutrients throughout the plant. Sap is distinct from latex, resin, or cell sap; it is a separ ...
. The birds keep the sap flowing apparently as a cavity defense mechanism against
rat snake Rat snakes are members – along with kingsnakes, milk snakes, vine snakes and indigo snakes – of the subfamily Colubrinae of the family Colubridae. They are medium to large constrictors and are found throughout much of the Northern Hemi ...
s and possibly other predators. The typical territory for a group ranges from about 125 to 200 acres (500,000 to 800,000 m²), but observers have reported territories running from a low of around 60 acres (240,000 m²), to an upper extreme of more than 600 acres (2.40 km²). The size of a particular territory is related to both habitat suitability and population density. Where red-cockaded woodpeckers occur at high densities, individuals appear to spend more time in territorial defense, potentially at the expense of foraging and time allocated to reproduction, resulting in reduced clutch size and fledgling production. The red-cockaded woodpecker plays a vital role in the intricate web of life of the southern pine forests. A number of other birds and small
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s use the cavities excavated by red-cockaded woodpeckers, such as
chickadee The chickadees are a group of North American birds in the tit family included in the genus '' Poecile''. Species found in North America are referred to as chickadees, while other species in the genus are called tits. They are small-sized bird ...
s,
bluebird The bluebirds are a North American group of medium-sized, mostly insectivorous or omnivorous birds in the order of Passerines in the genus ''Sialia'' of the thrush family (Turdidae). Bluebirds are one of the few thrush genera in the Americas. ...
s,
titmice ''Baeolophus'' is a genus of birds in the family Paridae. Its members are commonly known as titmice. All the species are native to North America. In the past, most authorities retained ''Baeolophus'' as a subgenus within the genus ''Parus'', b ...
, and several other woodpecker species, including the
downy Downy, also known as Lenor in Europe, Russia and Japan, is a brand of fabric softener produced by Procter & Gamble that was introduced in 1960. Lenor is a brand name of fabric softener and dryer sheets, also produced by P&G, sold in Europe, Rus ...
,
hairy Hairy may refer to: * people or animals covered in hairs or fur * plants covered in trichomes * insects covered in setae * people nicknamed "the Hairy" * Hairy (gene) See also * Hairies, a fictional people * Haerye ''Hunminjeongeum Haerye'' ...
, and
red-bellied woodpecker The red-bellied woodpecker (''Melanerpes carolinus'') is a medium-sized woodpecker of the family Picidae. It breeds mainly in the eastern United States, ranging as far south as Florida and as far north as Canada. Though it has a vivid orange-red ...
s. Larger woodpeckers such as northern flicker, red-bellied or
pileated woodpecker The pileated woodpecker (''Dryocopus pileatus'') is a large, mostly black woodpecker native to North America. An insectivore, it inhabits deciduous forests in eastern North America, the Great Lakes, the boreal forests of Canada, and parts of the ...
may take over a red-cockaded woodpecker cavity, sometimes enlarging the hole enough to allow
eastern screech owl The eastern screech owl (''Megascops asio'') or eastern screech-owl, is a small typical owl, owl that is relatively common in Eastern North America, from Mexico to Canada. This species is native to most wooded environments of its distribution, a ...
s,
wood duck The wood duck or Carolina duck (''Aix sponsa'') is a species of perching duck found in North America. The drake wood duck is one of the most colorful North American waterfowl. Description The wood duck is a medium-sized perching duck. A ty ...
s, and even
raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the common raccoon to distinguish it from other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of , and a body weight of ...
s to move in later. Flying squirrels, several species of
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s and
amphibia Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arbor ...
ns, and
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s, primarily bees and
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...
s, also will use red-cockaded woodpecker cavities.


Ecology

A significant competitor is the
pileated woodpecker The pileated woodpecker (''Dryocopus pileatus'') is a large, mostly black woodpecker native to North America. An insectivore, it inhabits deciduous forests in eastern North America, the Great Lakes, the boreal forests of Canada, and parts of the ...
, who is known for doubling the width of cavities in trees, making the tree as a whole uninhabitable to the red-cockaded woodpecker.


Conservation

The red-cockaded woodpecker suffers from habitat fragmentation when habitable pines are removed. When a larger cluster of birds gets split up, it is difficult for the young to find mates and eventually becomes an issue regarding species dispersal. While dispersing in search of new places to settle, the red-cockaded woodpecker encounters habitats of competing woodpecker species. The red-cockaded woodpecker has been the focus of conservation efforts even before the passing of the
Endangered Species Act The Endangered Species Act of 1973 (ESA or "The Act"; 16 U.S.C. § 1531 et seq.) is the primary law in the United States for protecting imperiled species. Designed to protect critically imperiled species from extinction as a "consequence of ec ...
in 1973. In Florida, pairs are being released at
DuPuis Management Area DuPuis Management Area is a 21,875 acre protected area in northwestern Palm Beach County, Florida and southwestern Martin County, Florida. Recreational opportunities include hunting, horseback riding, cycling, camping, hiking, auto touring, and fish ...
and other privately-owned land. Due to the high importance of nesting habitat on the woodpecker's reproduction, much management has been dedicated to create ideal and more numerous nesting sites. Nesting clusters have been spared from forestry activity to preserve old-growth, large diameter trees. The nesting sites themselves have also been managed to make them more appealing. The use of controlled burning has been used to reduce deciduous growth around nesting colonies. The red-cockaded woodpecker has been shown to prefer nesting sites with less deciduous growth. The use of controlled burning must be exercised with caution due to the highly flammable resin barriers formed by the woodpecker. In an effort to increase the red-cockaded woodpecker population, states such as Texas, Louisiana, Alabama, and Georgia's wildlife management are creating artificial cavities in Longleaf Pine trees. There are two methods in which wildlife management officers use to insert cavities in long leaf pines. The most respected and latest approach is to carve out a nesting cavity in the tree and insert a man-made rot-resistant wooden box with a PVC pipe small enough for only a red-cockaded woodpecker to fit through. These boxes, also known as "inserts", can last up to 10 years. The older and less used approach is to drill a cavity into the tree in hopes that the birds will settle there and nest. Due to the energetically expensive process of excavating new cavities, more energy is expended competing for existing home ranges rather than colonizing new areas. Cavities are highly sought-out resources by all cavity dwelling species and red-cockadeds have been observed roosting in them as early as the same night the boxes are installed. Alpha males usually reside in the best cavity, alpha females in the second best cavity until breeding season when their male partners allow their cavity to turn into the nest cavity. Juveniles are left with the lower quality cavities or no cavities at all, forcing them to roost on a branch outside overnight. Due to the fact that each family member requires a cavity to roost, land managers may choose to insert additional artificial cavities to boost survival of juveniles. Red-cockaded woodpeckers will even recolonize abandoned ranges when cavities are created. In addition to the creation of new cavities, methods for protecting existing cavities are also used. The most common technique employed is a restrictor plate. The plate prevents other species from enlarging or changing the shape of the cavity entrance. These restrictor plates must be carefully monitored, however, to ensure that no hindrance is given to the woodpecker. Adjustments must also be made as the tree grows. Southern flying squirrel exclusion devices may be considered as well. A study suggested that managers establish new woodpecker clusters away from streams and limit the use of excess cavities, both factors important in the recruitment of flying squirrels. Application of capsaicin on flying squirrel at cavities could be a cost-effective method.


Intentional habitat destruction by landowners

Private landowners are prohibited from modifying habitats or taking animals that are protected by the
Endangered Species Act The Endangered Species Act of 1973 (ESA or "The Act"; 16 U.S.C. § 1531 et seq.) is the primary law in the United States for protecting imperiled species. Designed to protect critically imperiled species from extinction as a "consequence of ec ...
(ESA). In response to the listing of the Red-cockaded woodpecker, it became common in Eastern Texas for landowners to begin cutting down the long-leaf pine trees so that the woodpeckers would not nest in their trees. By eliminating the trees on a piece of privately owned land, the land owners no longer have to abide by the requirements of the
U.S. Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with othe ...
biologists. By removing available habitat for the endangered species, landowners are able to benefit economically and maintain construction rights to the property. This led to the development of the Safe Harbor Agreements in which landowners agreed to manage their land in consistency with the conservation of a species of special conservation concern in return for relaxed ESA protection rules that allow a more flexible management of their own land.


See also

* Babbitt v. Sweet Home Chapter of Communities for a Great Oregon


References


Further reading

* *


External links


BirdLife Species Factsheet.The Nature Conservancy's Species page: Red Cockaded woodpecker
VIREO* ttp://ibc.hbw.com/ibc/phtml/especie.phtml?idEspecie=3880 Red-cockaded woodpecker videoson the Internet Bird Collection
A little bit more about the red-cockaded woodpecker and a few other endangered birds

Ecos.fws.gov
Species Profile
W.G.Jones State Forest, Conroe, TX
{{Authority control
red-cockaded woodpecker The red-cockaded woodpecker (''Leuconotopicus borealis'') is a woodpecker endemic to the southeastern United States. Description The red-cockaded woodpecker is small to mid-sized species, being intermediate in size between North America's two ...
Native birds of the Southeastern United States Endemic birds of the Eastern United States
red-cockaded woodpecker The red-cockaded woodpecker (''Leuconotopicus borealis'') is a woodpecker endemic to the southeastern United States. Description The red-cockaded woodpecker is small to mid-sized species, being intermediate in size between North America's two ...
red-cockaded woodpecker The red-cockaded woodpecker (''Leuconotopicus borealis'') is a woodpecker endemic to the southeastern United States. Description The red-cockaded woodpecker is small to mid-sized species, being intermediate in size between North America's two ...
ESA endangered species